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Nephrolepis cordifolia

(L.) C.Presl (1836)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Nephrolepidaceae Genus: Nephrolepis

kupukupu, kupukupu fern, kupukupu lau li‘i [kupukupu lau lii], narrow sword fern

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

  • Sometimes used on hula altars (kuahu) (Pukui 1942) and as a hula adornment (Valier 1995:74).

CULTURE

  • Keleanuinoho‘ana‘api‘api Lolale, chief of Lihue, grieves for his wife Kelea when she requests to leave to go sightseeing, "The fragrance of the kupukupu ferns, That are twisted about by the Waikoloa wind, By the Wai‘opua wind, My flower" (Kamakau 1991:47).

PROPAGATION/CULTIVATION

  • Grown in tissue culture; grows & spreads vigorously; new plants can be pulled out and replanted; thrive also in pots on lanais (Bornhorst 1996:72–73). Lighting conditions should be medium to high, with soil that is moist to dry (Hoshizaki & Moran 2002:391).

Natural History

Statewide Status

Indigenous

Island Status

Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Tent. Pterid.: 79 (1836)

Other References

Palmer 2003:189 (K, O, Mo, L, M, H); Staples & Herbst 2005:40 (DESCR, KEY); Hovenkamp & Miyamoto 2005:294 (DESCR, KEY); Fosberg 12437 BISH (L†, 1935)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Nephrolepis cordifolia Wahiawa Mt. Kauai Preserved_Specimen Forbes, C.N. 219 Kauai BISH 1907-08-01
2 Nephrolepis cordifolia Haalelepaakai Preserved_Specimen Munro, G.C. 168 Lanai BISH 1927-07-23
3 Nephrolepis cordifolia Kipapa Gulch, Waipio, south ridge, Koolau Mts. Preserved_Specimen Hosaka, E.Y. 736 Oahu BISH 1932-09-18
4 Nephrolepis cordifolia Lanipo Trail Preserved_Specimen Topping, D.L. 2675 Oahu BISH 1923-07-01
5 Nephrolepis cordifolia Koolau Mt. Punaluu Preserved_Specimen Rock, J.F.C. 219 Oahu BISH
6 Nephrolepis cordifolia Waianae Mts., ridge trail to Palikea Preserved_Specimen Cowan, R.S. 657 Oahu BISH 1947-07-02
7 Nephrolepis cordifolia Summit of Konahuanui Preserved_Specimen Forbes, C.N. 2191 Oahu BISH 1916-03-12
8 Nephrolepis cordifolia Waianae Mts.: Mauna Kapu - Palikea Preserved_Specimen Krajina, V.J. 611029015 Oahu BISH 1961-10-29
9 Nephrolepis cordifolia Power Line Trail between Hanalei and Lihue Preserved_Specimen Forbes, C.N. 153 Kauai BISH 1909-08-06
10 Nephrolepis cordifolia Honokahau Drainage Basin Preserved_Specimen Forbes, C.N. 511.511 Maui BISH 1917-09-25
11 Nephrolepis cordifolia East Maui, Upper Shelf of Kipahulu Valley, mauka of Ginger Camp Preserved_Specimen Welton, P. 1801 Maui BISH 1993-12-13
12 Nephrolepis cordifolia Main peak of Mr. Konahuanui Preserved_Specimen Garber, D.W. 319 Oahu BISH 1920-03-14
13 Nephrolepis cordifolia Kipahulu Valley, E part of valley, base of central pali Preserved_Specimen Lamoureux, C.H. 3921 Maui BISH 1967-08-07
14 Nephrolepis cordifolia Waikamoi, along lower flume Preserved_Specimen Hobdy, R.W. 392 Maui BISH 1978-11-15
15 Nephrolepis cordifolia Waialae Iki Trail Preserved_Specimen Woolford, H.A. 362 Oahu BISH 1956-01-01
16 Nephrolepis cordifolia Proposed Puu Ohu Reservoir, Kohala Mountains Preserved_Specimen Corn, C. Hawaii BISH 1984-03-27
17 Nephrolepis cordifolia Between Kalahuapueo and Hanalilolilo Preserved_Specimen Krajina, V.J. 620611097 Molokai BISH 1962-06-11
18 Nephrolepis cordifolia Kipahulu Valley, W part of valley Preserved_Specimen Lamoureux, C.H. 3954 Maui BISH 1967-08-09
19 Nephrolepis cordifolia West Maui, Lihue Preserved_Specimen Hobdy, R.W. 3300 Maui BISH 1991-06-04
20 Nephrolepis cordifolia Alakai Swamp, cloud zone. Preserved_Specimen Higgins, N.H. 95 Kauai BISH 1956-12-28
21 Nephrolepis cordifolia Forest of Waimea. Preserved_Specimen Lyons, Dr. Hawaii BISH
22 Nephrolepis cordifolia Hawaii Preserved_Specimen Morley, H. Hawaii BISH 1934-06-01
23 Nephrolepis cordifolia Kalanilehua, Olaa Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-16
24 Nephrolepis cordifolia Kilauea, Bird Park. Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-05
25 Nephrolepis cordifolia Makaha Valley Koolau Range. Preserved_Specimen Forbes, C.N. Oahu BISH 1909-02-12
26 Nephrolepis cordifolia Koolauloa Mts. between Punahua & Kaipaupau. Preserved_Specimen Forbes, C.N. Oahu BISH 1909-05-08
27 Nephrolepis cordifolia Kilauea, Bird Park Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-05
28 Nephrolepis cordifolia Kalanilehua, Olaa. Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-06-30
29 Nephrolepis cordifolia unknown Preserved_Specimen Lack, F. BISH 1880-01-01
30 Nephrolepis cordifolia W Maui, Puu Kukui, past cabin Preserved_Specimen Higashino, P.K. 1266 Maui BISH 1976-01-02
31 Nephrolepis cordifolia Konahuanui. Preserved_Specimen Lyon, H.L. Oahu BISH 1909-11-07
32 Nephrolepis cordifolia unknown: All the Islands. Preserved_Specimen BISH
33 Nephrolepis cordifolia Puna, Olaa FR, 10.8 km from jct. of Stainback and 17 Hwys moist, shady Preserved_Specimen Herat, T. 577 Hawaii BISH 1974-06-15
34 Nephrolepis cordifolia Waikolu Valley, steep valley wall below Puu Kaeo. Wet forest. Preserved_Specimen On ground. St.John, H. 23470 Molokai BISH 1948-12-22
35 Nephrolepis cordifolia SW flank of Mauna Loa, Manuka open Metrosideros forest on aa with Nephrolepis, Sadleria, Myrsine, Styphelia. Aa rocks covered with Cladina, Cladonia, Racomitrium Preserved_Specimen Higashino, P.K. 5985 Hawaii BISH 1977-07-03
36 Nephrolepis cordifolia Puu Kawiwi-Puu Kaala ridge. Waianae mts., Makaha- Waianae Kai moist forest Preserved_Specimen rhizome erect Fosberg, F.R. 10840 Oahu BISH 1935-03-31
37 Nephrolepis cordifolia Palikea, Southern Waianae Mountains. In side gully above Palawai gulch. Mixed native and alien forest. Plants terestrial but also epiphytic growing amongst Urera glabra, Asplenium acuminatum, Cibotium chammisoi, Metrosideros polymorpha, Acacia koa, Schinus terebinthifolia, Morella faya, Dicranopteris linearis, Cyrtandra waianaensis, and others. Preserved_Specimen Fronds narrow, pinnae gradually reducing towards base. Some tubers found, but these seem rare. Thomas, M.K. 127 Oahu BISH 2021-09-15
38 Nephrolepis cordifolia Kalaheo Container grown plant from residence of J.Hipp. Preserved_Specimen Also known as N. cordifolia Flynn, T.W. 2768 Kauai BISH 1988-02-23
39 Nephrolepis cordifolia N of Saddle Rd in kipuka, closed Metrosideros forest, with shrubs Preserved_Specimen Wagner, W.L. 4695 Hawaii BISH 1982-11-18
40 Nephrolepis cordifolia Honolulu, Hawaii Herbarium of the H.L. Lyon Arboretum Preserved_Specimen Fern to 0.5m. Frond to 0.6m. Stem Brown. Stem and rachis lightly tomentose. Midrib of pinnae black. Spores brown. No. 2. Teraoka, W. 31 Oahu BISH 1979-06-03
41 Nephrolepis cordifolia Kona-Hilo Hwy. ~6.8 mi past Hongo Store; up unpaved road ~100 m; ~20 m NE of road Metrosideros-Cibotium forest Preserved_Specimen ~1' fern; epiphyte Newell, C.L. 1019 Hawaii BISH 1967-06-26
42 Nephrolepis cordifolia Kalani Prison Rd. ~7.4 mi from intersection with Kona-Hilo Hwy.; ~50 m into foresgt at L of road Metrosideros, Cibotium, Pritchardia Preserved_Specimen ~2' fern; epiphyte Newell, C.L. 346 Hawaii BISH 1966-08-09
43 Nephrolepis cordifolia Poamoho trail; Paalaa Wahiawa Koolau Range On ground, side of trail Preserved_Specimen Wilson, K.A. 130 Oahu BISH 1952-04-06
44 Nephrolepis cordifolia Top of cliffs at head of Haiku Valley. Main divide of Koolau Mts. Heeia. Wet exposed ridge Preserved_Specimen rhizome erect Fosberg, F.R. 13870 Oahu BISH 1937-05-23
45 Nephrolepis cordifolia Head of Waikolu Valley. Hanalilolilo,. Rocky slopes with thin soil, above edge of pali; open low forest. Preserved_Specimen St.John, H. 12356 Molokai BISH 1932-12-21
46 Nephrolepis cordifolia Kilauea area, near crater Open, modified site; lava cracks. Preserved_Specimen Fern w. wiry thizomes, lacking tubers; fronds erect. Sykes, W.R. 34591 Hawaii BISH 1991-06-02
47 Nephrolepis cordifolia East Maui, Hana District, S fork of Waiohonu ridge above stream bank, on Metrosideros Preserved_Specimen Epiphyte. Harrison, B. 18 Maui BISH 1972-06-14
48 Nephrolepis cordifolia 2nd Gulch east of Puu Kaupakuhale, Mokuleia, Puu Kaala, Wainae Mts. wooded ridge E. of gulch Preserved_Specimen Fosberg, F.R. 8914 Oahu BISH 1932-10-23
49 Nephrolepis cordifolia Kona-Hilo Hwy. ~6.8 mi past Hongo Store; up unpaved road ~100 m; ~20 m NE of road Metrosideros-Cibotium forest Preserved_Specimen ~6 fern; epiphyte Newell, C.L. 1018 Hawaii BISH 1967-06-26
50 Nephrolepis cordifolia S. Hilo, L branch off Saddle Rd., ~26 km from Hilo in shade, among grasses Preserved_Specimen ~1/3 m high fronds Herat, T. 740 Hawaii BISH 1974-06-22
51 Nephrolepis cordifolia South Hilo distr., Hilo, Walmart garden center, 325 Makaala St. Potted plants for sale in shade area. Preserved_Specimen Fronds stiffly erect, pinnules dark green, round, with white dots around margin on upper side. Common name: lemon buttons Staples, G.W. 1283 Hawaii BISH 2007-02-15
52 Nephrolepis cordifolia Kona, Keauhou No. 1; new Bishop Estate road. ohia rain forest Preserved_Specimen Christ, J.H. Hawaii BISH 1958-05-15
53 Nephrolepis cordifolia W slope of Hualalai, above Kailua, Kona edge of lava gulch Preserved_Specimen Gillett, G.W. 1705 Hawaii BISH 1965-01-21
54 Nephrolepis cordifolia Fern Estates Rd. ~1.1 mi.; ~25 m E of road Metrosideros-Cibotium forest Preserved_Specimen ~1' fern; epiphyte Newell, C.L. 628 Hawaii BISH 1966-08-28
55 Nephrolepis cordifolia Main divide, crest of Koolau Mts., near head of Castle Trail, Kaipaupau wet forest Preserved_Specimen fronds erect. Pinnae slightly curved dorsally Fosberg, F.R. 13954 Oahu BISH 1937-05-31
56 Nephrolepis cordifolia SW flank of Mauna Loa, Manuka closed Metrosideros forest on aa with Cibotium, Nephrolepis, Ilex, Myrsine, Pipturus, Psychotria, Vaccinium Preserved_Specimen Higashino, P.K. 5867 Hawaii BISH 1977-07-20
57 Nephrolepis cordifolia Poamoho summit in gullly. Terrestrial, growing in thich bryophye mat of Bazzania cordistipula, Dicranodontium porodictyon, and Pagiochila deflexa. Also growing with Nephrolepis exaltata, Dicranopteris linearis, Cibotium menziesii, and Sadleria pallida in Metrosideros dominated wet forest. Preserved_Specimen Tubers not present in specimen, but were present in small amounts along the roots. Thomas, M.K. 102 Oahu BISH 2021-08-18
58 Nephrolepis cordifolia Bishop Museum campus next to back entrance to Hawaii Hall. Cultivated in landscape with Phymatosorus grossus and Microlepia srigosa. Preserved_Specimen Blades with overlapping pinnae bases on abaxial surface. Pinnae almost perpendicular to rachis. Thomas, M.K. 130 Oahu BISH 2021-09-21
59 Nephrolepis cordifolia Aukele Street, Kailua, Oahu, Hawaii Cultivated, growing in crushed lava rock, wet conditions. Preserved_Specimen Sword fern, clumping, sending out runners with tubers. Tuber with long soft hairs. Crago, L.M. 2005229 Oahu BISH 2005-11-14
60 Nephrolepis cordifolia Top of cliff at head of Wailau Valley. Rain forest. Preserved_Specimen Fronds more or less erect, others erect to recumbent. Fosberg, F.R. 13368 Molokai BISH 1936-12-23
61 Nephrolepis cordifolia East Maui, Koolau Gap, Waikamoi Preserve. USFWS Tr#4, below lower fence. Koolau District Disturbed Wet Lowland Forest single fertile frond Preserved_Specimen Welton, P. 2052 Maui BISH 1998-05-29
62 Nephrolepis cordifolia Puu Peahinaia. Thick woods. Preserved_Specimen Topping, D.L. 3321 Oahu BISH 1929-04-28
63 Nephrolepis cordifolia East Maui, Waihoi Valley on tree branches Preserved_Specimen Epiphyte. Harrison, B. 6 Maui BISH 1972-09-21
64 Nephrolepis cordifolia Olokui Wailau, ridge between upper fork's of Wailele Stream. Moist forest. Preserved_Specimen St.John, H. 23322 Molokai BISH 1948-02-06
65 Nephrolepis cordifolia Pupukea-Kahuku Trail. Moist bank. Preserved_Specimen Topping, D.L. 3331 Oahu BISH 1929-05-12
66 Nephrolepis cordifolia Main divide, crest of Koolau Mts., between Waikane Valley and Waikakalaua Gulch Wet windswept ridge. Preserved_Specimen fronds erect, pinnae slightly curved dorsally. Fosberg, F.R. 13982 Oahu BISH 1937-05-30
67 Nephrolepis cordifolia Kona, Keauhou, above Kahaluu FR, ~4.7 mi above lower gate in jeep road and wet Metrosideros forest Preserved_Specimen Herbst, D.R. 5409 Hawaii BISH 1975-07-15
68 Nephrolepis cordifolia Honolulu, 42 Coelho Way In shade, under Musa, Heliconia humilis, H. metallica, and Phaeomeria magnifica. Preserved_Specimen Perrenial fern; never fruits. Neal, M.C. 1247 Oahu BISH 1944-12-02
69 Nephrolepis cordifolia Waimea district; Waiakoali, Na Pali Kona Forest Reserve Metrosideros-Cheirodendron montane wet slopes and drainage with Syzygium sandiwcensis, Kadua affinis, Pittosporum gayanum, P. glabrum, Ilex anomala, Melicope anisata, M. clusiifolia, Myrsine dentata, M. punctata, Psychotria hexandra, P. mariniana, P. greenwel Preserved_Specimen terrestrial rhizome short erect, growing in small colonies, common Wood, K.R. 15218 Kauai BISH 2012-09-14
70 Nephrolepis cordifolia Nuuanu in forest east of Pali lookout parking lot. Southern Koolau mountains. Growing on log in Cinnamomum dominated forest. Also growing with Asplenium nidus, Casuarina, and Microsorum grossum. Preserved_Specimen Fronds narrowly lanceolate. Pinnae nearly perpendicular to rachis.On undersurface, base of pinnae overlapping rachis and auriculate. Small tubers found but were rotten. Thomas, M.K. 266 Oahu BISH 2022-02-21
71 Nephrolepis cordifolia Palikea Suspected naturalized growing in large clump. Area was formerly managed by TNC. Maybe outplanted? Preserved_Specimen Garrett, N. 575 Oahu BISH 2023-03-09
72 Nephrolepis cordifolia Mt. Kaala. Waianae District Metrosideros-Rugosa forest Preserved_Specimen Grant, M.L. 7463 Oahu BISH 1934-10-04
73 Nephrolepis cordifolia Kaala. Along FAA road overlooking Makaleha valley near three points. Mesic ridge community with Dicranopteris linearis, Pteridium aqulinum subsp. decompositum, Metrosideros polymorpha, Nephrolepis exaltata, Leptecophylla tameameae, Schinus terebinthifolius, Psidium cattleianum, Mellinis repens, and others. Preserved_Specimen Lance shaped fronds on creeping rhizome. Pinnae falcate and coriaceous in sunlight. Shaded fronds longer and pinnae less coriaceous. Pinnae bases truncate and slightly overlapping rachis in sunny fronds. Shaded plants with pinnae bases not overlapping rachis. Thomas, M.K. 462 Oahu BISH 2022-10-20
74 Nephrolepis cordifolia Wahiawa, central ridge to Puu Kaala Growing among lobeliaceous plants Preserved_Specimen Chock, A.K. 469 Oahu BISH 1952-05-25
75 Nephrolepis cordifolia East Maui, Koolau Gap, Waikamoi Preserve, USFWS TR#4. Between stations 90-92. Koolau District. Disturbed Wet Lowland Forest Preserved_Specimen 7 fronds with rhizome, some fertile Welton, W. 2049 Maui BISH 1998-05-28
76 Nephrolepis cordifolia Puu Hapapa, Central Waianae Mountains. Growing on a sunny slope next to the snail enclosure perimeter and next to fence line. Forest type = mesic/dry, mostly invaded with Schinus, Toona, and Psidium with some native elements. Preserved_Specimen Plants match the cultivated type of Nephrolepis cordifolia seen at nurseries. Plants with pinnae almost perpendicular to the plane of the blade. Pinnae narrow with basal portions overlapping rachin on undersurface. Base of pinnules auricled even when sterile. Tubers abundant. Thomas, M.K. 341 Oahu BISH 2022-04-13
77 Nephrolepis cordifolia McCall's Flower Farm. Haunani rd. Volcano, Hawaii. Cultivated from wild material in nearby forest. Preserved_Specimen Narrowly lanceolate pinnate ferns. Pinnae lanceolate, bases auriculate and overlapping rachis on abaxial surface. Lynch, K. Hawaii BISH 2022-02-23
78 Nephrolepis cordifolia East Maui, Makawao District, Makawao Forest Reserve Terrestrial, locally common in dense shade under Eucalyptus, Fraxinus, Acaia, Metrosideros Preserved_Specimen Oppenheimer, H.L. 92128 Maui BISH 2021-09-28
79 Nephrolepis cordifolia West Maui, Wailuku District, slopes of Hanaula; three ridges south of Waikapu Valley. On steep degraded slope. Vegetation: Metrosideros, Dicranopteris, Freycinetia, Myrsine, Pritchardia, Vaccinium, Erigeron, Lythrum, Blechnum, Adiatum. Preserved_Specimen Erect narrow, terrestrial fern, unusual form more often observed in cultivation than in the wild. Oppenheimer, H.L. 31309 Maui BISH 2013-03-13
80 Nephrolepis cordifolia Olaa FR, trail off back road with Metrosideros collina, Cheirodendron trigynum, Cibotium glaucum, Cyrtandra, Peperomia, Dryopteris Preserved_Specimen Higashino, P.K. 7186 Hawaii BISH 1978-03-23
81 Nephrolepis cordifolia E rim of Ukumehame Gulch open moist Metrosideros forest Preserved_Specimen St.John, H. 25656 Maui BISH 1955-12-28
82 Nephrolepis cordifolia Puu Konahuanui. Tree trunks. Preserved_Specimen Topping, D.L. 3211 Oahu BISH 1926-02-07
83 Nephrolepis cordifolia Southeast slope of Kaala, Oahu, T.H. Naturalized and common in forest near top. Preserved_Specimen Degener, O. 12238 Oahu BISH
84 Nephrolepis cordifolia Laie Trail, Kahawainui. Ihiihi Ridge, Koolau Mts., Laie wet forest Preserved_Specimen epiphytic Fosberg, F.R. 14235 Oahu BISH 1937-07-24
85 Nephrolepis cordifolia Alakai Swamp trail From the forest immediately above the 2nd bog, growing on the ground Preserved_Specimen Takeuchi, W.N. 80 Kauai BISH 1982-11-13
86 Nephrolepis cordifolia Lanaihale, ridge moist forest Preserved_Specimen Fosberg, F.R. 12437 Lanai BISH 1935-11-29
87 Nephrolepis cordifolia SW flank of Mauna Loa, Manuka closed Metrosideros and Acacia on weathered aa; with Cibotium, Myoporum, Myrsine, Vaccinium Preserved_Specimen Higashino, P.K. 6371 Hawaii BISH 1977-08-07
88 Nephrolepis cordifolia Main divide, crest of Koolau Mts. above Kaipaupua Gulch edge of small open bog Preserved_Specimen pinnae somewhat backward curved Fosberg, F.R. 14227 Oahu BISH 1937-07-24
89 Nephrolepis cordifolia Waianae Mts., Pule'e Gulch. Growing epiphytically and terrestrially in Schinus terebinthifolius and Psidium guajava-dominated habitat. Preserved_Specimen Looks like cultivated type of N. cordifolia. Elevation very low for occurrence of Hawaiian N. cordifolia. A few dozen individuals. Kawelo, K. 542 Oahu BISH 2020-11-15
90 Nephrolepis cordifolia Volcano: Hwy. 11, 22.5 miles from Hilo. Akatsuka Orchid Gardens In planter bed. Preserved_Specimen Wilson, K.A. 2440 Hawaii BISH 1994-05-31
91 Nephrolepis cordifolia Lyon Arboretum, Ethnobotany garden Growing as epiphytes on cultivated Metrosideros polymorpha and on ground. Preserved_Specimen Dakine get plenty tubers! Thomas, M.K. 132 Oahu BISH 2021-10-14
92 Nephrolepis cordifolia upper Olaa FR, back road on tree fern trunks, dense stand of Cibotium Preserved_Specimen fronds pendent, flat, no tubers on rhizomes Fosberg, F.R. 51738 Hawaii BISH 1969-12-09
93 Nephrolepis cordifolia Honolulu, Hawaii Herbarium of the H.L. Lyon Arboretum Preserved_Specimen Fern to 0.5m. Frond to 0.6m. Stem Brown. Stem and rachis lightly tomentose. Midrib of pinnae black. Spores brown. No. 2. Teraoka, W. 30 Oahu BISH 1979-06-03
94 Nephrolepis cordifolia Kaipapau. Northern Koolau Mountains On cut bank above stream bed in a hanging valley near the Koolau summit. Area dominated by mostly native wet forest of Metrosideros polymorpha, Polyscias oahuensis, Cheirodendron, Psychotria, Pritchardia, Dicranopteris, Macherina, Sadleria, and others. Plants growing with Amauropelta globulifera, Isachne pallens, Diplazium sandwichianum, Carex alligata, Cyrtandra paludosa, and others. Preserved_Specimen Erect fern with arching fronds. Tubers present in colony (one present on specimen). Fronds narrowly lanceolate, pinnae slightly falcate with rruncate bases. Thomas, M.K. 447 Oahu BISH 2023-09-14
95 Nephrolepis cordifolia Olaa FR, Wright Rd., Puna Cibotium glaucum and Metrosideros forest Preserved_Specimen St.John, H. 25070 Hawaii BISH 1953-01-29
96 Nephrolepis cordifolia Wailuku River, 12 mi. W of Hilo on tree trunks or mossy ground; Metrosideros, semi-open, mossy Preserved_Specimen Fosberg, F.R. 60587 Hawaii BISH 1980-09-08
97 Nephrolepis cordifolia On Waianae range. On summit of Mt. Kaala In somewhat open area Preserved_Specimen With long thin fronds Herat, T. 527 Oahu BISH 1973-09-30
98 Nephrolepis cordifolia Poamoho summit near cabin Epiphytic on base of Metrosideros polymorpha. Growing in a thick bryophyte community dominated by Bazzania cordistipula. Preserved_Specimen Plant with narrow fronds and blunt, falcate pinnae. Tubers present. Thomas, M.K. 101 Oahu BISH 2021-08-17
99 Nephrolepis cordifolia Fern Estates Rd. ~1.1 mi.; ~25 m E of road Metrosideros-Cibotium forest Preserved_Specimen ~1' fern; epiphyte Newell, C.L. 649 Hawaii BISH 1966-08-28
100 Nephrolepis cordifolia East Maui, Waihoi Valley old log with Polypodium and Trachypodopsis Preserved_Specimen Harrison, B. 2596 Maui BISH 1972-11-21
101 Nephrolepis cordifolia Main divide, crest of Koolau Mts., Uncle Tom's Cabin, Peak at head of Waiawa trail Wet windswept ridge. Preserved_Specimen Fronds erect. Pinnae slightly curved back. Fosberg, F.R. 13912 Oahu BISH 1937-05-30
102 Nephrolepis cordifolia Waipio, above Kipapa Gulch, summit of Koolau mts. wet forest Preserved_Specimen Fosberg, F.R. 8674 Oahu BISH 1932-09-18
103 Nephrolepis cordifolia Lanihuli, summit Growing on rotten wood Preserved_Specimen MacDaniels, L.H. 167 Oahu BISH 1927-01-23
104 Nephrolepis cordifolia Kalaupapa; Papapaiki Preserved_Specimen medium terrestrial fern Hughes, G.D. 1117 Molokai BISH 2004-12-09
105 Nephrolepis cordifolia Chain of Craters, Napau Crater Trail Preserved_Specimen Stone, B.C. 3059 Hawaii BISH 1959-08-07
106 Nephrolepis cordifolia Olinda area Maui. Olinda flume (4260 ft) then down to 3000 feet level and across road across Waikamoi (3000 ft) stream, up opposite bank to top (3075 ft) then down road to tree. Preserved_Specimen Tree loaded with Elaphoglossum, microdenium. Gillett, G.W. 1771 Maui BISH 1965-06-16
107 Nephrolepis cordifolia Olokele Ditch Trail, Makaweli, Preserved_Specimen St.John, H. 23083 Kauai BISH 1947-12-26
108 Nephrolepis cordifolia Kalaupapa; Waikolu, upper valley floor, above intake tunnel Preserved_Specimen medium terrestrial fern Wysong, M.L. 728 Molokai BISH 2005-03-29
109 Nephrolepis cordifolia East Maui, Between Kaumakani fence tag 30-31, 200m below camp. Preserved_Specimen Common name: Boston fern, sword fern, tuber ladd Welton, P. 2179 Maui BISH 2004-02-24
110 Nephrolepis cordifolia Schofield Barracks, South Range Preserved_Specimen Mesic forest. Patch 20 m x 20 m. Kawelo, K. 100 Oahu BISH 2008-09-18
111 Nephrolepis cordifolia Alakai Swamp Trail In the forest adjoining the 1st bog Preserved_Specimen Takeuchi, W.N. 11 Kauai BISH 1982-08-01
112 Nephrolepis cordifolia Pauoa-Pacific Heights ridge. Preserved_Specimen Fern specimens Garber, D.W. 346 Oahu BISH 1920-03-21
113 Nephrolepis cordifolia Puu Lanahuli Preserved_Specimen Topping, D.L. 2726 Oahu BISH 1923-04-29
114 Nephrolepis cordifolia Kaholuamanu Preserved_Specimen Rock, J.F.C. 1469 Kauai BISH 1909-03-03
115 Nephrolepis cordifolia W Maui, Kaulalewelewe, Puu Kukui 19.9N 156.6W Preserved_Specimen Krajina, V.J. 620418006 Maui BISH 1962-04-18
116 Nephrolepis cordifolia Paalaa, Wahiawa, Koolau Mountains Dry forest Preserved_Specimen Higgins, V.H. 42 Oahu BISH 1955-11-20
117 Nephrolepis cordifolia East Maui, Nahiku, Kuhiwa trail, Koolau Forest Reserve wet woods, on mossy trunks by stream Preserved_Specimen St.John, H. 17920 Maui BISH 1937-01-02
118 Nephrolepis cordifolia North Halawa-Heeia Divide, Haiku Valley. Koolau Range, Cloud Zone in low moist thicket on ridge. Preserved_Specimen St.John, H. 20427 Oahu BISH 1943-12-12
119 Nephrolepis cordifolia Alakai Swamp trail, above 2nd bog forest Preserved_Specimen Takeuchi, W.N. 79 Kauai BISH 1982-11-13
120 Nephrolepis cordifolia Waimea Dist.: Kokee State park, Na Pali-Kona Forest Reserve; Alakai Swamp trail from parking area at trail head to Kawaikoi. Stream crossing; ca. 1/4 mi beyond jct. betw Alakai and Pihea trails Preserved_Specimen Terrestrial; stolons long-creeping, forming white tubers sporatically; fronds erect to spreading Wagner, W.L. 6423 Kauai BISH 1990-06-21
121 Nephrolepis cordifolia Preserved_Specimen BISH
122 Nephrolepis cordifolia unknown Preserved_Specimen BISH
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